Citation Rankings
Highly Cited Researchers

University of Denver Highly Cited Researchers CitationNatural Sciences / Biological Science Rankings 2025

The Citation Rankings (Highly Cited Researchers) highlight the most cited and influential researchers across a vast range of scientific fields, covering 197 main and sub-disciplines, including the Art and Humanities Citation Rankings and Social Sciences and Humanities Citation Rankings. This service, offered exclusively by AD Scientific Index, aims to provide a balanced perspective by excluding data from CERN and other statistical sources that disproportionately favor certain researchers, particularly in the social sciences and humanities. This unique approach addresses the advantages researchers with multi-author publications and CERN involvement hold over others. Furthermore, citation rankings help uncover rule violations and unethical practices within the research community. For more detailed citation analyses, explore the Scientists Last 6 Years Citation Index, Universities Total Citations Rankings 2025, and Universities Last 6 Years Citations Rankings 2025.

* Total Citation Counts Rankings
Ranking Based On Selection :1
United States
Paul C Sutton
Ecological Economics
Sustainability Science
Population Geography
Urban Science
Global
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
61,388
27,809
0.453
* Total Citation Counts Rankings
Ranking Based On Selection :2
United States
Craig T Jordan
cancer
leukemia
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
30,285
12,546
0.414
* Total Citation Counts Rankings
Ranking Based On Selection :3
United States
Sara Branco
ecology
evolution
fungi
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
7,622
5,914
0.776
* Total Citation Counts Rankings
Ranking Based On Selection :4
United States
Diana Tomback
Forest ecology
conservation biology
applied ecology
plant-animal interactions
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
6,829
1,731
0.253
* Total Citation Counts Rankings
Ranking Based On Selection :5
United States
Martin Margittai
Alzheimer's Disease
Protein Biochemistry
Neurodegeneration
Structural Biology
Protein Aggregation
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
5,095
1,438
0.282
* Total Citation Counts Rankings
Ranking Based On Selection :6
United States
Ryan C Hill
Proteomics of the ECM
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
4,991
4,168
0.835
* Total Citation Counts Rankings
Ranking Based On Selection :7
United States
John G Swallow
Evolutionary Physiology
Animal Behavior
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
4,873
1,059
0.217
* Total Citation Counts Rankings
Ranking Based On Selection :8
United States
Ilene Grabel
Developmental Architectures
Global Financial Governance
Int'l Financial Institutions
Financial Crises
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
4,280
1,072
0.250
* Total Citation Counts Rankings
Ranking Based On Selection :9
United States
Greg Cronin
Applied Ecology
Transdisciplinary Research
Ecosystem Restoration
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,994
809
0.203
* Total Citation Counts Rankings
Ranking Based On Selection :10
United States
Michael Wunder
Animal Population Ecology
Animal Behavior
Conservation Biology
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,425
1,755
0.512
* Total Citation Counts Rankings
Ranking Based On Selection :11
United States
Kristin Artinger
Developmental Biology
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,398
1,114
0.328
* Total Citation Counts Rankings
Ranking Based On Selection :12
United States
Kingshuk Ghosh
Proteins
Polymers
Biophysics
Statistical Physics
Data Science
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,369
1,755
0.521
* Total Citation Counts Rankings
Ranking Based On Selection :13
United States
Christy Briles
Paleoecology and Biogeography
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,311
1,563
0.472
* Total Citation Counts Rankings
Ranking Based On Selection :14
United States
Scott A Nichols
animal origins
evolutionary biology
cell biology
sponges
immunity
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,240
1,030
0.318
* Total Citation Counts Rankings
Ranking Based On Selection :15
United States
Patrick H Martin
forest ecology
invasion ecology
tropical montane forest ecology
disturbance ecology
global change
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,171
1,567
0.494
* Total Citation Counts Rankings
Ranking Based On Selection :16
United States
Brian Buma
Landscape Ecology
Disturbance
Resilience
Succession
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,991
2,420
0.809
* Total Citation Counts Rankings
Ranking Based On Selection :17
United States
Alan M Vajda
Endocrine Disruption
Ecological Developmental Biology
Conservation Biology
Applied Ecology
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,901
1,176
0.405
* Total Citation Counts Rankings
Ranking Based On Selection :18
United States
Caley Orr
paleoanthropology
primate functional morphology
human evolution
hand
wrist
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,804
1,374
0.490
* Total Citation Counts Rankings
Ranking Based On Selection :19
United States
Kristofer S Fritz
Proteomics
Sirtuins
Acetylation
Alcohol-associated Liver Disease
Diabetic Nephropathy
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,775
1,585
0.571
* Total Citation Counts Rankings
Ranking Based On Selection :20
United States
Timberley Roane
Environmental Microbiology
Microbial Ecology
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,672
985
0.369
* Total Citation Counts Rankings
Ranking Based On Selection :21
United States
Annika C Mosier
microbial ecology
microbial biogeochemistry
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,477
948
0.383
* Total Citation Counts Rankings
Ranking Based On Selection :22
United States
JC Fogleman
Cytochrome P450
forensic genetics
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,326
356
0.153
* Total Citation Counts Rankings
Ranking Based On Selection :23
United States
Joseph Angleson
Cell Biology
Ca2+ and cAMP signaling
GPCRs
Melanocortin Receptors
regulated exocytosis
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,278
283
0.124
* Total Citation Counts Rankings
Ranking Based On Selection :24
United States
Robin Tinghitella
Behavioral ecology
sexual selection
parental effects
speciation
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,269
1,319
0.581
* Total Citation Counts Rankings
Ranking Based On Selection :25
United States
Michael J Greene
Sociobiology
Behavioral Ecology
Chemical Ecology
Comparative Physiology
Animal Behavior
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,184
616
0.282
* Total Citation Counts Rankings
Rankings
7,164
United States
3,390
2
Ranking Based On Selection: 1
Paul C Sutton
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
61,388
27,809
0.453
Ecological Economics
Sustainability Science
Population Geography
Urban Science
Global
* Total Citation Counts Rankings
Rankings
21,910
United States
10,114
30
Ranking Based On Selection: 2
Craig T Jordan
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
30,285
12,546
0.414
* Total Citation Counts Rankings
Rankings
133,106
United States
50,886
161
Ranking Based On Selection: 3
Sara Branco
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
7,622
5,914
0.776
* Total Citation Counts Rankings
Rankings
149,600
United States
56,194
180
Ranking Based On Selection: 4
Diana Tomback
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
6,829
1,731
0.253
Forest ecology
conservation biology
applied ecology
plant-animal interactions
* Total Citation Counts Rankings
Rankings
200,444
United States
71,862
52
Ranking Based On Selection: 5
Martin Margittai
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
5,095
1,438
0.282
Alzheimer's Disease
Protein Biochemistry
Neurodegeneration
Structural Biology
Protein Aggregation
* Total Citation Counts Rankings
Rankings
204,534
United States
73,097
245
Ranking Based On Selection: 6
Ryan C Hill
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
4,991
4,168
0.835
* Total Citation Counts Rankings
Rankings
209,371
United States
74,562
249
Ranking Based On Selection: 7
John G Swallow
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
4,873
1,059
0.217
* Total Citation Counts Rankings
Rankings
236,413
United States
82,485
62
Ranking Based On Selection: 8
Ilene Grabel
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
4,280
1,072
0.250
Developmental Architectures
Global Financial Governance
Int'l Financial Institutions
Financial Crises
* Total Citation Counts Rankings
Rankings
252,011
United States
87,065
288
Ranking Based On Selection: 9
Greg Cronin
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,994
809
0.203
Applied Ecology
Transdisciplinary Research
Ecosystem Restoration
* Total Citation Counts Rankings
Rankings
287,890
United States
97,403
319
Ranking Based On Selection: 10
Michael Wunder
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,425
1,755
0.512
Animal Population Ecology
Animal Behavior
Conservation Biology
* Total Citation Counts Rankings
Rankings
289,872
United States
97,971
322
Ranking Based On Selection: 11
Kristin Artinger
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,398
1,114
0.328
* Total Citation Counts Rankings
Rankings
292,024
United States
98,571
83
Ranking Based On Selection: 12
Kingshuk Ghosh
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,369
1,755
0.521
Proteins
Polymers
Biophysics
Statistical Physics
Data Science
* Total Citation Counts Rankings
Rankings
296,458
United States
99,823
331
Ranking Based On Selection: 13
Christy Briles
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,311
1,563
0.472
* Total Citation Counts Rankings
Rankings
301,911
United States
101,366
86
Ranking Based On Selection: 14
Scott A Nichols
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,240
1,030
0.318
animal origins
evolutionary biology
cell biology
sponges
immunity
* Total Citation Counts Rankings
Rankings
307,289
United States
102,908
89
Ranking Based On Selection: 15
Patrick H Martin
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
3,171
1,567
0.494
forest ecology
invasion ecology
tropical montane forest ecology
disturbance ecology
global change
* Total Citation Counts Rankings
Rankings
322,525
United States
107,170
347
Ranking Based On Selection: 16
Brian Buma
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,991
2,420
0.809
Landscape Ecology
Disturbance
Resilience
Succession
* Total Citation Counts Rankings
Rankings
330,765
United States
109,519
358
Ranking Based On Selection: 17
Alan M Vajda
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,901
1,176
0.405
Endocrine Disruption
Ecological Developmental Biology
Conservation Biology
Applied Ecology
* Total Citation Counts Rankings
Rankings
339,982
United States
112,037
369
Ranking Based On Selection: 18
Caley Orr
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,804
1,374
0.490
paleoanthropology
primate functional morphology
human evolution
hand
wrist
* Total Citation Counts Rankings
Rankings
342,866
United States
112,853
374
Ranking Based On Selection: 19
Kristofer S Fritz
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,775
1,585
0.571
Proteomics
Sirtuins
Acetylation
Alcohol-associated Liver Disease
Diabetic Nephropathy
* Total Citation Counts Rankings
Rankings
353,441
United States
115,749
382
Ranking Based On Selection: 20
Timberley Roane
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,672
985
0.369
Environmental Microbiology
Microbial Ecology
* Total Citation Counts Rankings
Rankings
375,061
United States
121,766
400
Ranking Based On Selection: 21
Annika C Mosier
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,477
948
0.383
* Total Citation Counts Rankings
Rankings
393,456
United States
126,754
111
Ranking Based On Selection: 22
JC Fogleman
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,326
356
0.153
* Total Citation Counts Rankings
Rankings
399,678
United States
128,416
115
Ranking Based On Selection: 23
Joseph Angleson
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,278
283
0.124
Cell Biology
Ca2+ and cAMP signaling
GPCRs
Melanocortin Receptors
regulated exocytosis
* Total Citation Counts Rankings
Rankings
400,716
United States
128,679
116
Ranking Based On Selection: 24
Robin Tinghitella
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,269
1,319
0.581
Behavioral ecology
sexual selection
parental effects
speciation
* Total Citation Counts Rankings
Rankings
412,175
United States
131,840
424
Ranking Based On Selection: 25
Michael J Greene
Citation Metrics
Total
Last 6 Years
Last 6 Years / Total
2,184
616
0.282
Sociobiology
Behavioral Ecology
Chemical Ecology
Comparative Physiology
Animal Behavior